Prof SM Dippenaar


Research Interests


I am interested in the biodiversity, distribution, systematics and evolution of freshwater free-living copepods as well as symbiotic Siphonostomatoida occurring on vertebrate hosts. This includes the population genetics of species occurring on the same or different host species of elasmobranchs.

Popular articles

George W. Benz & Susan M. Dippenaar . 2000. Putting the Bite on Jaws: Copepods as Enemies of Sharks. The Shark Tagger 1998 Summary. Newsletter of the NOAA: 12,18.

D. Engelbrecht, J. Grosel & S. Dippenaar. 2007. The western population of Short-clawed Lark Certhilauda chuana in South Africa revisited. Bulletin of the African Bird Club 14: 58-61.

Susan Dippenaar. 2009. Hooked on the Unknown. Beyond Blue, April 2009, Issue 3: 28-32.

Susan Dippenaar. 2010. Symbiotic marine copepods. Friendz, October 2010, Issue 10: 26.


DIPPENAAR, S.M. & FERGUSON, J.W.H. 1994. Towards a captive breeding programme for the Riverine Rabbit Bunolagus monticularis . South African Journal of Science 90: 381 - 385.

DIPPENAAR, S.M. & FERGUSON, J.W.H. 1994. Use of DNA fingerprinting in planning a breeding programme for the riverine rabbit ( Bunolagus monticularis ). Zoo Biology 13(3): 231 - 243.

OLIVIER, P.A.S., DIPPENAAR, S.M. & VAN NIEKERK, J.P. 1997. A first report on the morphology of the postantennal process in Lernanthropus (Lernanthropidae: Copepoda) and its possible significance as a taxonomic feature. South African Journal of Zoology 32(2):43-48.

DIPPENAAR, S.M. & OLIVIER, P.A.S. 1999. New morphological information of the parasitic copepod Kroyeria dispar Wilson , 1935 (Copepoda: Kroyeriidae) from the east coast of South Africa . South African Journal of Zoology 34(3):125-129.

OLIVIER, P.A.S., DIPPENAAR, S.M. , KHALIL, L.F. & MOKGALONG , N.M. 2000. Observations on a lesser-known monogenean, Udonella myliobati , from a copepod parasite, Lepeophtheirus natalensis , parasitising the spotted ragged-tooth shark, Carcharias taurus , from South African waters. Onderstepoort Journal of Veterinary Research 67:135-140.

DIPPENAAR, S.M. , BENZ, G.W. & OLIVIER, P.A.S. 2000. Kroyeria deetsi n.sp. (Kroyeriidae: Siphonostomatoida), a parasitic copepod infecting gills of spinner sharks, Carcharhinus brevipinna (Mùller & Henle, 1839), in the Indian Ocean . African Zoology 35:185-192.

DIPPENAAR, S.M. , OLIVIER, P.A.S. & BENZ, G.W. 2001. Kroyeria sphyrnae Rangnekar, 1957 (Copepoda, Siphonostomatoida, Kroyeriidae): first description of the male, supplementary remarks on the female, a new geographic record for the species, and a key to Kroyeria males. Crustaceana 74(9):883-894.

DIPPENAAR, S.M. , LUUS-POWELL, W.J. & ROUX, F. 2001. Lamproglena hoi n.sp. (Copepoda: Lernaeidae) from two yellowfish hosts, Barbus marequensis and Barbus polylepis , caught in a river in Mpumalanga , South Africa . Onderstepoort Journal of Veterinary Research 68:209-215.

BULLARD , S.A. & DIPPENAAR, S.M . 2003. Branchotenthes robinoverstreeti n. gen. and n. sp. (Monogenea: Hexabothriidae) from gill filaments of the bowmouth guitarfish, Rhina ancylostoma (Rhynchobatidae), in the Indian Ocean . Journal of Parasitology 89(3):595-601.

BULLARD , S.A. , DIPPENAAR, S.M. , HOFFMAYER, E.R. & BENZ, G.W. 2004. New locality records for Dermophthirius carcharhini Mac Callum, 1926 (Monogenea: Microbothriidae) and D. maccallumi Watson and Thorson, 1976 and a list of hosts and localities for species of Dermophthirius Mac Callum, 1926. Comparative Parasitology 71(1): 78-80.

DIPPENAAR, S.M. , OLIVIER, P.A.S. & BENZ, G.W. 2004. Schistobrachia jordaanae n. sp. (Copepoda: Siphonostomatoida: Lernaeopodidae) from gill filaments of a diamond ray (G ymnura natalensis ) captured in the Indian Ocean , and a key to species of Schistobrachia , Dendrapta, and B rianella . Journal of Parasitology 90: 481-484.

DIPPENAAR, S.M. 2004. Reported siphonostomatoid copepods parasitic on marine fishes of southern Africa . Crustaceana 77(11): 1281-1328.

ENGELBRECHT, G.D. & DIPPENAAR, S.M. 2005. Nestling growth, plumage development and rate of prey delivery at a nest of the Desert Cisticola, Cisticola aridula . African Zoology 40: 151-153.

MODIBA, M.A., DIPPENAAR, S.M. & DIPPENAAR-SCHOEMAN, A.S. 2005. A checklist of spiders from Sovenga Hill, an inselberg in the Savanna Biome, Limpopo Province , South Africa (Arachnida: Araneae). Koedoe 48(2): 109-115.

KHOZA, T.T., DIPPENAAR, S.M. & DIPPENAAR-SCHOEMAN, A.S. 2005. The biodiversity and species composition of the spider community of Marion Island , a recent survey (Arachnida: Araneae). Koedoe 48(2): 103-107.

DIPPENAAR, S.M. & JORDAAN, B.P. 2006. Nesippus orientalis Heller, 1868 (Pandaridae: Siphonostomatoida): descriptions of the adult, young and immature females, a first description of the male and aspects of their functional morphology. Systematic Parasitology 65: 27-41.

DIPPENAAR, S.M. & JORDAAN, B.P. 2007. New host and geographical records of siphonostomatoid copepods associated with elasmobranchs off the KwaZulu-Natal coast, South Africa . Onderstepoort Journal of Veterinary Research 74(2): 169-175.

DIPPENAAR, S.M. & JORDAAN, B.P. 2008. Description of the adult female and male of Naobranchia kabatana n.sp. (Siphonostomatoida: Lernaeopodidae) from Muraenesox bagio (Hamilton) (Muraenesocidae) caught in the Indian Ocean, South Africa . Systematic Parasitology 70: 27-34.

DIPPENAAR, S.M. , DIPPENAAR-SCHOEMAN, A.S., MODIBA, M.A. & KHOZA, T.T. 2008. A checklist of the spiders (Arachnida, Araneae) of the Polokwane Nature Reserve, Limpopo Province , South Africa . Koedoe 50(1): 10-17.

DIPPENAAR, S.M. , VAN TONDER, R.C., WINTNER, S.P. & ZUNGU, P. 2008. Spatial distribution of Nemesis lamna Risso, 1826 (Copepoda: Siphonostomatoida: Eudactylinidae) on the gills of white sharks Carcharodon carcharias off Kwa-Zulu Natal, South Africa. African Journal of Marine Science 30(1): 143-148.

DIPPENAAR, S.M., VAN TONDER, R.C. & WINTNER, S.P. 2009. Is there evidence of niche restriction in the spatial distribution of Kroyeria dispar Wilson, 1935, K. papillipes Wilson, 1932 and Eudactylina pusilla Cressey, 1967 (Copepoda: Siphonostomatoida) on the gill filaments of tiger sharks Galeocerdo cuvier off KwasZulu-Natal, South Africa? Hydrobiologia 619:89-101.

DIPPENAAR, S.M. 2009. Estimated molecular phylogenetic relationships of six siphonostomatoid families symbiotic on elasmobranchs. Crustaceana 82(12): 1547-1567.

DIPPENAAR, S.M., MATHIBELA, R.B. & BLOOMER, P. 2010. Cytochrome oxidase I sequences reveal possible cryptic diversity in the cosmopolitan symbiotic copepod Nesippus orientalis Heller, 1868 (Pandaridae: Siphonostomatoida) on elasmobranch hosts from the KwaZulu-Natal coast of South Africa. Experimental Parasitology 125: 42-50.

DIPPENAAR, S.M. & JORDAAN, B.P. 2012. Notes on the morphology and ecology of the adult females of Nesippus species (Siphonostomatoida: Pandaridae) with a key for identification. Zootaxa 3170: 18-30.

DIPPENAAR, S.M. 2012. A redescription of Pseudocharopinus pteromylaei Raibaut et Essafi, 1979 (Siphonostomatoida: Lernaeopodidae) collected from the South African east coast. Folia Parasitologica 59(3): 216-220.

DIPPENAAR, S.M. & LEBEPE, M.C. 2013. Two new species of Pupulina (Caligidae, Siphonostomatoida) from mobulid rays (Mobulidae) off South Africa. Systematic Parasitology 85: 27-35. DOI:10.1007/s11230-013-9406-y

MAAKE, P.A., MWALE, M., DIPPENAAR, S.M. & GON, O. 2013. Mitochondrial and nuclear DNA reveals a complete lineage sorting of Glossogobius callidus (Teleostei: Gobiidae) in southern Africa. African Journal of Aquatic Science 38 (suppl. 1): 15-29. DOI: 10.2989/16085914.2013.782263

LEBEPE, M.C. & DIPPENAAR, S.M. 2013. A report of symbiotic Siphonostomatoida (Copepoda) infecting mobulids (Rajiformes: Mobulidae) off the KwaZulu-Natal coast, South Africa. African Zoology 48(2): 326-332.

MANGENA, T., JORDAAN, B.P. & DIPPENAAR, S.M. 2014. Phylogenetic relationships and genetic diversity of Nemesis Risso, 1826 species found on different elasmobranch host species, off the KwaZulu-Natal coast. African Journal of Marine Science 36(2): 163-173.

LUCAS, K.C., DIPPENAAR, S.M. & DU TOIT, P.H. 2014. Analysis of assessment practice and subsequent performance of third year level students in natural sciences. African Education Review 11(4): 563-583.

DIPPENAAR, S.M. & JORDAAN, A. 2015. How Achtheinus spp. (Pandaridae: Siphonostomatoida) females attach to their elasmobranch hosts with notes on the effects on the host. Folia Parasitologica 62: 005. doi: 10.14411/fp.2015.005

MOKUMO, P.J. & DIPPENAAR, S.M. 2015. Reports of Kroyeria species collected from South African waters with notes on the host-parasite associations. African Zoology 50(2):127-132.

DIPPENAAR, S.M. & MOLELE, R.A. 2015. Siphonostomatoid copepods infecting Squalus acutipinnis Regan, 1908 off South Africa. African Journal of Marine Science 37:605-608. DOI:

LEBEPE, M.C. & DIPPENAAR, S.M. 2016. Barnard’s Brachiella sp., Parabrachiella supplicans (Barnard, 1955) and Eubrachiella sublobulata (Barnard, 1955) (Copepoda: Siphonostomatoida: Lernaeopodidae) deposited in the Iziko South African Museum. Zootaxa 4061(1): 051-060.

DIPPENAAR, S.M. 2016. Does Alella Leigh-Sharpe, 1925 (Siphonostomatoida: Lernaeopodidae) really consists of seven species? Systematic Parasitology 93:47-56. DOI 10.1007/s11230-015-9601-0

DIPPENAAR, S.M. 2016. Biodiversity and studies of marine symbiotic siphonostomatoids off South Africa. African Journal of Marine Science 38:1-5. DOI

DIPPENAAR, S.M. 2016. Schistobrachia kabata sp. nov. (Siphonostomatoida: Lernaeopodidae) from Rajiform hosts off South Africa. Zootaxa 4147:104-113. DOI

DIPPENAAR, S.M. 2017. Trebius benzi n. sp. (Siphonostomatoida: Trebiidae) infecting Squalus acutipinnis Regan off South Africa. Systematic Parasitology 94:91-96. DOI

DIPPENAAR, S.M. 2018. Symbiotic Siphonostomatoida (Copepoda) collected from white sharks, Carcharodon carcharias (Lamniformes, Lamnidae), during the OCEARCH expedition along the coast of South Africa. Crustaceana 91(1):103-111. DOI 10.1163/15685403-00003746

DIPPENAAR, S.M. 2018. Description of four new species and a revision of the genus Tripaphylus Richiardi in Anonymous, 1878 (Copepoda: Siphonostomatoida: Sphyriidae). Systematic Parasitology 95:173-200. DOI: 10.1007/s11230-017-9767-8

DIPPENAAR, S.M. 2018. Resurrection of Alebion difficile (van Beneden, 1892) (Copepoda: Siphonostomatoida: Caligidae) with notes on other species of Alebion Krøyer, 1863 collected from elasmobranchs off South Africa and an updated key of the adult females. Systematic Parasitology. DOI: 10.1007/s11230-018-9811-3